Project Summary
Karen Wishner
U.S. GLOBEC: Cross-frontal Distributions and Exchange of
Zooplankton on Georges Bank
ABSTRACT
Cross-frontal distributions and exchange of zooplankton on Georges Bank
are a primary focus of Phase 3 of the U.S. GLOBEC Georges Bank program,
which has the broader goal of understanding the response of marine
populations to global climate change, especially interactions between
zooplankton and fish. This proposal examines the cross-frontal exchange of
zooplankton at two sites important to the biology of GLOBEC-targeted
species: the Northeast Peak Front in March (coincident with expected inflow
of the copepod Calanus finmarchicus and the presence of cod larvae and high
predator abundance) and the Southern Flank Tidal Mixing Front in June
(coincident with high concentrations of Calanus and predators). Questions
addressed include the interaction between vertically-stratified zooplankton
distributions and the cross-frontal exchange at different depths, the temporal
variability of these interactions on diel, tidal, and fortnightly scales, and the
effects over time of the repeated passage of tidal fronts at particular locations
along a bathymetric slope. Zooplankton abundances and vertical distributions
will be determined from a Lagrangian sampling scheme utilizing transects
relative to frontal position conducted at different phases of the tidal cycle
(semidiurnal and fortnightly), and will be related to hydrographic structure,
tidal phase, and distance from the front. In a complementary Eulerian
sampling scheme employing time series sampling at fixed locations along a
bathymetric slope, zooplankton distributions and variability will be related to
the time since frontal passage. Vertically-stratified MOCNESS tows will be
used to collect zooplankton. Cross-frontal exchange of zooplankton (different
species and lifestages with different vertical distributions) will be estimated
from concentration gradients across the front at different depths and along
isopycnal surfaces, using various estimates of flow and water exchange
provided by physical oceanographers.
BACKGROUND
This proposal, on the cross-frontal distributions and exchange of
zooplankton on Georges Bank, is one of 3 linked proposals submitted
for GLOBEC Phase III and recommended for funding. The other 2
proposals in the group are (1) Hebert / Barth and (2) Gifford /
Sieracki / Belkin. Hebert / Barth is a physical oceanography
proposal while the Gifford et al. and Wishner proposals combine
biology and physics.
The major scientific questions addressed by the three proposals
as a whole are:
- What is the 3-dimensional circulation associated with the target
fronts over a fortnightly time-scale? (Hebert and Barth;
Belkin)
- What is the effect of tidal current variability over
semi-diurnal and fortnightly time-scales? (Hebert and Barth;
Belkin)
- What are the dynamics and rates of cross-frontal exchange of
water, including the cross-frontal fluxes and turbulent mixing
rates? (Hebert and Barth; Belkin)
- What water masses are present? (Hebert and Barth; Belkin)
- What are the distributions of target zooplankton species across
the fronts? (Wishner)
- What is the potential prey field (nanoplankton and
microplankton) of target copepods and and larval fish in the frontal
regions? (Gifford and Sieracki)
- What are the rates of feeding and growth of Calanus
nauplii and copepodids at different cross-frontal locations?
(Gifford and Sieracki)
- What are the cross-frontal exchange rates of zooplankton
(Wishner), nano- and microplankton? (Gifford and
Sieracki)
- How do biological and physical processes interact to control
cross-frontal exchange of target organisms?
(Wishner; Gifford, Sieracki and Belkin). For
example, how do vertical distributions and migration of zooplankton
interact with cross-frontal exchange processes at different depths
and frontal locations? (Wishner)
- How does the spatial variability of frontal movement influence
the distribution and exchange of organisms at different locations
and depths? (Wishner; Gifford, Sieracki and
Belkin)
- How do temporal variations of the front (e.g., spring/neap,
tidal, and diel scales) interact with populations of target
organisms? (Wishner; Gifford, Sieracki and Belkin)
- What are the potential consequences of these processes for
recruitment of target organisms? (Wishner;
Gifford, Sieracki and Belkin)
The work will be done as a coordinated two-ship operation, using
one ship (Oceanus) for the SEASOAR work of Hebert and Barth and one
ship (Endeavor) for the biological/physical work of Wishner and
Gifford, Sieracki, and Belkin. Cross-frontal processes and exchange
will be studied at two sites and times important to the biology of
GLOBEC-targetted species and known to have strong thermal fronts at
those times: the Northeast Peak Front in March (coincident with
expected inflow of the copepod Calanus finmarchicus and the
presence of cod larvae and high predator abundance) and the Southern
Flank Tidal Mixing Front in June (coincident with high
concentrations of Calanus and predators). Two 18-day
cruises have been requested on the Endeavor by Wishner and Gifford
et al., and a 16 and 18 day cruise have been requested on the
Oceanus by Hebert / Barth.
Specific cruise activities, which would be done in each area,
include:
- High resolution SEASOAR surveys to map the frontal region
(radiator pattern) and shorter surveys (butterfly pattern) around an
isopycnal float deployed, tracked, and recovered several times in
the frontal areas (Hebert and Barth)
- High resolution SCANFISH surveys in the frontal areas
(Belkin)
- Cross-frontal transects of zooplankton
(Wishner, MOC-1, vertically-stratified tows),
microplankton, and nanoplankton (Gifford, Sieracki, CTD, GO-FLO
rosette), and hydrography (Belkin, CTD, SCANFISH)
- Time series stations (same instruments as above)
(Wishner; Gifford, Sieracki and Belkin)
- Zooplankton vital rates (Gifford and Sieracki)
Sensors mounted on the SEASOAR include dual temperature and
conductivity sensors, a fluorometer, transmissometer, and a
Microsoar. Sensors mounted on the SCANFISH include temperature and
conductivity sensors, a fluorometer, and oxygen sensor. MOCNESS
sensors include temperature, salinity, a fluorometer, and
transmissometer. The SEASOAR (Hebert and Barth) and SCANFISH
(Belkin in conjunction with biologists) will be doing near real-time
mapping of cross-frontal distributions of a wide variety of
physical, biological, and chemical parameters, which will be used to
determine cross-frontal exchanges and mixing of water. These data
will be used in conjunction with the species distributional data
from the MOCNESS and CTD-Go-FLO rosette to determine cross-frontal
exchanges of nanophytoplankton, nanozooplankton, microphytoplankton,
microzooplankton, and mesozooplankton. Thus, zooplankton
distributions and variability will be analyzed within the context of
a sophisticated physical understanding of cross-frontal exchange and
dynamics. Our sampling scheme takes account of the semi-weekly and
diurnal tidal cycle, day/night cycle, front location, and Bank
topography.
Other direct collaborators, who will participate in these
cruises, include Barbara Sullivan (cross-frontal distribution and
exchange of small predators in MOC-1 samples), Larry Madin (predator
distributions and feeding rates), and Ann Bucklin
(Pseudocalanus cross-frontal distributions and genetics from
MOC-1 samples). We will also coordinate closely with the mooring
work (Bob Beardsley: Northeast Peak moorings, Jim Irish: Southern
Flank moorings), remote sensing (Jim Bisagni: front detection), and
shipboard ADCP analyses (Charlie Flagg), as well as other
biologists, physicists, and modelers.