Three Main Biological Effects of MPAs
 
“Reserve Effects” = higher density in closed areas, demographic shifts, reproductive output increased, potentially resulting in:
“Spill Over” = net movement out of adults (apart from ontogenetic or other non-density related movements)
A Really Big Question: Is there a “Fatal Attraction” of fishing effort to reserve boundaries, seeking spillover?
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“ Export” = net export of larvae or juveniles to open areas
Fisheries benefit from permanently closed areas via Spillover, Export, or overall reductions in fishing rate