Calanus IBM overview
Egg
N1-2
N3
N4-6
C1-3
C4
C5
C5
Female
diapause
DVM
Non-feeding
Foraging
Behavior
The model groupings
Growth rate, G:
G = gmax * Food / (ks + Food)
Where, gmax = b + m*ln (T)
b and m are empirically fit*
Development rate, R:
R = Ω* a (T + b) -2.05
a and b are empirically fit
Where Ω is a penalty
function that decreases development
rate at extremely low food levels
Mortality:
Each stage has a fixed mortality
rate (decreasing
logarithmically by stage)
Copepods also die if their age
within stage is greater
than that predicted at their minimum allowable
T
*Data
for empirically fit parameters from Campbell et al. (2001), Vidal (1980),
and/or Peterson (1986),
depending on species