Calanus IBM overview
Growth rate, G:
G = gmax * Food / (ks + Food)
Where, gmax = b + m*ln (T)
b and m are empirically fit*
Development rate, R:
R = Ω* a (T + b) -2.05
a and b are empirically fit
Where Ω is a penalty function that decreases
development rate at extremely low food levels
Mortality:
Each stage has a fixed mortality rate
(decreasing logarithmically by stage)
Copepods also die if their age within stage is
greater than that predicted at their minimum
allowable T
The model groupings
C5
C5
C4
C1-3
N4-6
N3
N1-2
*Data for empirically fit parameters from Campbell et al. (2001), Vidal (1980), and/or
Peterson (1986), depending on species