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Growth rate, G:
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G = gmax * Food /
(ks + Food)
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Where, gmax = b +
m*ln (T)
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b and m are
empirically fit*
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Development
rate, R:
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R = Ω* a (T
+ b) -2.05
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a and b are
empirically fit
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Where Ω is
a penalty function that decreases
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development rate
at extremely low food levels
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Mortality:
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Each stage has a
fixed mortality rate
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(decreasing
logarithmically by stage)
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Copepods also
die if their age within stage is
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greater than
that predicted at their minimum
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allowable T
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