S1: Uptake (
P
on
N
)
S1=
μ
max
f(I)f(N)f(T)P
S2: Grazing (
Z
on
P
)
S2=
g
max
f(P)f(T)Z
S3: Loss of
P
S3=λ
P
S4: Remineral. of
D
S4=ε
D
S5: Loss of
Z
S5=m
Z
2
Copepod Population
4-Stage (ENCA)
Lower Food Web
Eggs
S5
Z
γS5
Nauplii
α
S2
Copepodids
S2
βS2
P
Adults
S3
S1
N
D
S4