S1: Uptake (P on N)
S1=μmaxf(I)f(N)f(T)P
S2: Grazing (Z on P)
S2=gmaxf(P)f(T)Z
S3: Loss of P
S3=λ P
S4: Remineral. of D
S4=εD
S5: Loss of Z
S5=mZ2
Copepod Population
Lower Food Web
Eggs
S5
µZ
γS5
Nauplii
αS2
Copepodids
S2
βS2
P
Adults
S3
S1
N
D
S4